There was/is a thread running in this forum from an OP who noted that one side of his system was noticeably stronger than the other side and he was looking for troubleshooting tips.
As far as your tubes, I don’t have any experience with your gear, but I do have a test CD that has a specific tone for testing AC voltage (with a multimeter set to ACV) at the speaker terminals or amp terminals. I have noted that the tubes in the balanced input sockets of my preamp will have an effect on thie readings. In other words, if one side has a higher reading than the other side, if I swap the balanced input tubes around, the high and low readings follow the tubes.
Where I was going with all of that, is maybe you have a weak tube(s) in your tube complement and it isn’t your hearing causing the issue you are experiencing.
(As far as your power tubes, I would have thought that if the bias was equal on both sides, that would compensate for a weak tube. After re-reading your thread, maybe I was wrong in my thinking.)