Definition 1.5 was a livelier speaker than Def2, and its tonal center of gravity was shifted toward the upper midrange compared to Def2. The downside of Def1.5 was the MDF structure and the "cabinet talk" resulting from it at high SPLs. With Def2, Zu dramatically reduced the cabinet talk of the Definition architecture, giving it even more useful dynamic range, but one of the costs of tuning out the glare in Def1.5 was the slightly overdamped sound of Def2 (only in comparison to the incredibly jumpy Def1.5). The supertweeter network was improved as well, to tune out some of the older speaker's false sparkle evident in the supertweeter's anomalies. Def2 went as high, but it just didn't have all the tickle that Def1.5 had, along with that speaker's very top end distortions.
Def4 resolves the difference completely. Def3 should mostly. All of the liveliness of Def1.5 is restored in Def4, and then some. And all of the discipline and accuracy of Def2 is retained, and then some. The speed and openness of the nano-FRDs coupled to the tonal illumination provided by the smoothly extended Radian supertweeter eliminates the dark-tilt evident in Def2.
Solid State amps on Def2 went a long way to lifting the trace darkness in its tone, as did objective tube amps with extended top ends and bursty dynamic traits, but now that character isn't just mitigated. It's gone from the Def4 design and should be very much reduced in Def3's Def2 roots as well, since the nano-drivers are significantly responsible for the change (not the Radian alone). Def2s upgraded with 2010 HO FRDs pretty much lost the Def2 dark tilt anyway, so consider the small Def2 "overcorrection" to be adjusted out regardless which upgrade path you take.
Phil