I'd say that pretty much everything across the board has improved. I'm having to go by memory with the DAC2 since it has been discontinued for quite a while so I haven't played with one recently. So, I definitely can't give a detailed comparison. From memory, it has higher resolution than the DAC2 while remaining very smooth and unfatiguing. Tonally it is balanced more neutrally than the DAC2 as well. I personally found the DAC2 to be slightly overly warm and the DAC3 isn't. Don't misinterpret that as meaning that the DAC3 is cool, because it isn't at all IMHO. Considering that it is almost twice the price of the DAC2 it is no surprise that it is a considerably improved piece.
Sorry, I don't know the answer to your USB question.