Again, thank you guys for the contributions. Please keep sending along any thoughts you have. So far they are exactly the kind of answers I was hoping for from current owners. Wish I could invite you all over for a drink and a listen!
So I played around for a few hours tonight and managed to improve the bass presentation on the speakers significantly. I've had to find my own settings, as the examples posted above just haven't done it for me, in my space, with my tastes. I would still say that these MkIV's in their current configuration here are outputting only maybe 70% of the bass quality I was getting from the Mk1.5's. But steps forward for sure.
I've definitely had this realization.... I lived (very happily!) with my previous Zu's for 12 years here and had them dialled in just perfectly. I probably had somewhere around 5000 hours of listening bliss with them. Then I found these used MkIV's locally for a good price and decided to make the leap. Somehow I thought I could just slot them in just where the others were and figured it would be instant love. Now I get it... they are a different speaker, with a different solution for transducing bass frequencies, and I'm going to have to play around and get to know them in time...
I was struck by reading this thread --
https://forum.audiogon.com/discussions/zu-definition-4-height -- where 52tiger says he "was running out of options and really wasn't sure if the Def 4 was better than the def3," but after raising the height off the floor "the bass is tight and amazing, the downfiring design is so much better than the rear firing its no comparison." Clearly I have some experimenting to do.
I'm having faith in the last line of that thread by dentdog: "With my Def 4s I worked them around quite a bit to optimize... For me it was well worth all the effort."