John, when I was purchasing my Io, I asked Jim White what differences he heard between the standard Io and Io Signature, and with and without the second power supply option. His reply as to what he heard was very direct:
The Signature provides the following benefits:
.. Greater resolution
.. More neutral presentation
.. Improved high frequency extension and detail
.. Greater top to bottom coherency
.. Better bass definition
Adding the second supply improves the sound in the following ways:
.. Added space
.. Improved macro dynamics
.. Improved micro dynamics
.. Blacker backround
.. Greater ease to the sound, less strained