The box has to hold the body of the speaker stable so that the energy generated by the amplifier to move the cone is converted only into motion of the cone. This is never perfectly achieved, of course.The answer to your question is in the "this is never perfectly achieved"...
There is some critical value where resonance and distortions increase, and a treshold under which the distortions and resonance dont implicate an audible effects...
Putting springs under speakers isolate so much well from the other speakers influence and from feed back effect from the floor and from other external vibrations, then the audible effect is beneficial because the movement of the cone is maintained under a critical threshold...The isolation from external influence is less detrimental to the cone movements than some internal resonance from the speakers box without isolation coming from these external factors ,especially from the other twin speaker in the room...
It is not the case when speakers are not isolated properly.... It is the reason why springs are very useful.... Before i use springs i was using my sandwiches of different materials, that was good damping and complementary coupling and decoupling but not so powerful radical decoupling than springs.....The audible effect was way better than using nothing, but a smearing of the sound subsisted that was erased by the presence of the springs....